The Mocalw Bear Spirit bulldozed the Gluttony Termites by charging over them with its huge size. Every now and then the mandibles of the Gluttony Termites pierced into the bottom of the Mocalw Bear's paws as it stomped over them but that could hardly be considered an issue.
After all, the Mocalw Bear Spirit's body would heal as long as it was in possession of soul particles, and it was not as if the mandibles could inflict any damage other than a few scratches to the Mocalw Bear Spirit, in the first place.
The bulldozing didn't kill every single Gluttony Termite but it caused enough havoc to affect their entire colony. Not knowing whom to attack first and who to focus on, the Gluttony Termites threw their heads left and right, attacking blindly. It took them a while to find their target, and most of them were already dead or crippled by then.